Bryan D. Devan, Ph.D.
Associate Professor
Chief, Laboratory of Comparative Neuropsychology (LCN)
Psychology Department
Towson University

Office: LA3146
Phone: 410-704-3727

Mailing Address:

Psychology Department
Towson University
8000 York Rd.
Towson, MD  21252


Research Methods in Psychology PSYC.314
Physiological Psychology PSYC.465
Advanced Biological Psychology PSYC.674
Directed Research in Psychology (PSYC-391)
Independent Investigation in Psychology (PSYC-491)

Pub Links:

background pubs
LCN pubs



Devan B.D., Nufar C., Piscopello J. Deibel S.L., & McDonald, R.J. (2015) Cognitive and Stimulus-Response Habit Functions of the Neo- (Dorsal) Striatum. In Vinoth, J. Series: Innovations in Cognitive Neuroscience. The Basal Ganglia-Novel Perspectives on Motor and Cognitive Functions. submitted for publication.

Devan B.D., Magalis C and McDonald R.J. (2015). Hippocampal development and the dissociation of cognitive-spatial mapping from motor performance [version 2; referees: 2 approved] F1000Research, 4:625 (doi: 10.12688/f1000research.6966.2)

Devan, B.D., Pistell, P.J., Duffy, K.A., Kelley-Bell, B., Spangler, E.L., & Ingram, D.K. (2014). Phosphodiesterase inhibition facilitates cognitive restoration in animal models of age-related memory decline. NeuroRehabilitation, 34, 101-111.

Devan, B.D., Hong, N.S., McDonald, R.J. (2011). Parallel Associative Processing in the Dorsal Striatum: Segregation of Stimulus-Response and Cognitive Control Subregions. Neurobiology of Learning and Memory, 96, 95-120.

Turchi, J., Devan, B., Pingbo, Y., Sigrist, E., & Mishkin, M. (2010). Pharmacological evidence that both cognitive memory and habit formation contribute to within-session learning of concurrent visual discriminations. Neuropsychologia, 48, 2245-2250.

Pistell, P.J., Nelson, C.M., Miller, M.G., Ingram, D.K, Devan, B.D. (2009). Striatal lesions interfere with acquisition of a complex maze task in rats. Behavioural Brain Research, 197(1):138-43.

Duffy, K.B., Spangler, E.L. Devan, B.D., Guo, Z.H., Bowker, J.L., Janas, A.M. Hagepanos, A., Minor, R.K., de Cabo, R., Mouton, P.R., Shukitt-Hale, B., Joseph, J.A., Ingram, D.K. (2008) A blueberry-enriched diet provides cellular protection against oxidative stress and attenuates a kainate-induced learning impairment in rats. Neurobiology of Aging, 29(11):1680-9.

Pistell, P.J., Daffin Jr., L.W., Nelson, C.M., Duffy, K.B., Bowker, J.L., Spangler, E.L., Ingram, D.K., Devan, B.D. (2007). Combined administration of subthreshold doses of the nitric oxide inhibitor, nitro-L-arginine, and muscarinic receptor antagonist, scopolamine, impairs complex maze learning in rats. Behavioural Pharmacology, 18, 801-805.

Devan, B.D., Pistell, P.J., Daffin Jr., L.W., Nelson, C.M., Duffy, K.B., Bowker, J.L., Bharati, I.S., Sierra-Mercado Jr., D., Spangler, E.L., & Ingram, D.K. (2007). Sildenafil citrate attenuates a complex maze impairment induced by intracerebroventricular infusion of the NOS inhibitor Nω-nitro-L-arginine methyl ester. European Journal of Pharmacology, 563(1-3),134-140.

Devan, B.D., Bowker, J.L., Duffy, K.B., Bharati, I.S., Jimenez, M, Sierra-Mercado Jr., D., Nelson, C.M., Spangler, E.L., & Ingram, D.K. (2006). The phosphodiesterase inhibitor sildenafil citrate attenuates the learning impairment in rats induced by nitric oxide synthase inhibition. Psychopharmacology, 183, 439-445.

Devan, B.D., Bowker, J.L., Bharati, I.S., Duffy, K.B., Nelson, C.M., Daffin Jr., L.W., Spangler, E.L., & Ingram, D.K. (2005). Phosphodiesterase type 5 (PDE5) inhibition and cognitive enhancement. Drugs of the Future, 30(7), 725-736.

Janas, A., Cunningham C.V., Duffy, K.B., Devan, B.D., Grieg N., Hollaway H., Yu Q-S., Markowska, A., Ingram D.K., & Spangler, E. L. (2005). The cholinesterase inhibitor, phenserine, improves Morris water maze performance of scopolamine-treated rats. Life Sciences, 76, 1073-1081.

Devan, B.D., Sierra-Mercado Jr., D., Jimenez, M., Bowker, J.L., Duffy, K.B., Spangler, E.L., & Ingram, D.K. (2004). Phosphodiesterase inhibition by sildenafil citrate attenuates the learning impairment induced by blockade of cholinergic muscarinic receptors in rats. Pharmacology, Biochemistry and Behavior, 79, 691-699.

McDonald, R.J., Hong, N.S., & Devan, B.D. (2004). The challenges of understanding mammalian cognition and memory-based behaviors: An interactive learning and memory systems approach. Neuroscience & Biobehavioral Reviews, 28, 719-745.

McDonald, R. J., Devan, B. D., & Hong, N. S. (2004). Multiple memory systems: The power of interactions. Neurobiology of Learning and Memory, 82, 333-346.

Stouffer, E.M., Petri, H.L., & Devan, B.D. (2004). Effect of D-serine on delayed match-to-place task for the water maze. Behavioural Brain Research, 152, 447-452.

Devan, B.D., Stouffer, E.M., Petri, H.L., McDonald, R.J., & Olds, J.L. (2003). Partial reinforcement across trials impairs escape performance but spares place learning in the water maze. Behavioural Brain Research, 141, 91-104.

Devan, B.D., Petri, H.L., Mishkin, M., Stouffer, E.M., Bowker, J.L., Yin, P.B., Buffalari, D.M., & Olds, J.L. (2002). A room with a view and a polarizing cue: Individual differences in the stimulus control of place navigation and passive latent learning in the water maze. Neurobiology of Learning and Memory, 78, 79-99.

Devan, B. D., & McDonald, R. J. (2001). A cautionary note on interpreting the effects of partial reinforcement on place learning performance in the water maze.  Behavioural Brain Research, 119, 213-216.

Devan, B. D., Goad, E.H., Petri, H.L., Antoniadis, E.A., Hong, N. S., Ko, C.L., Leblanc, L., Lebovic, S.S., Lo,Q., Ralph, M.L., & McDonald, R.J. (2001). Circadian phase-shifted rats show normal acquisition but impaired long term retention of place memory in the water task. Neurobiology of Learning and Memory, 75, 51-62.

Devan, B.D., & White, N.M. (1999). Parallel information processing in the dorsal striatum: Relation to hippocampal function. Journal of Neuroscience, 19, 2789-2798.

Devan, B.D., McDonald, R.J., & White, N.M. (1999). Effects of medial and lateral caudate-putamen lesions on place- and cue-guided behaviors in the water maze: Relation to thigmotaxis. Behavioural Brain Research, 100, 5-14.

Devan, B. D., Goad, E.H., & Petri, H.L.(1996). Dissociation of hippocampal and striatal contributions to spatial navigation in the water maze. Neurobiology of Learning and Memory, 66, 305-323.

Devan, B. D., Blank, G.S., & Petri, H.L. (1992). Place navigation in the Morris water task: Effects of reduced platform interval lighting and pseudorandom platform positioning. Psychobiology, 20, 120-126.